Activities

Eve Marder

Variability, Homeostasis, and Compensation in Rhythmic Neuronal Networks

Neurons can live many years while ion channels and receptors turn over in neuronal membranes in minutes, hours, days or weeks. Therefore homeostatic mechanisms are needed to provide stable neuronal and network function over an animal's lifetime. Consequently, it is important to determine how tightly regulated synaptic and intrinsic properties must be to ensure adequate network performance. Moreover, it is important to determine the extent to which compensatory mechanisms allow for multiple solutions to the production of similar behavior. I will use examples from theoretical and experimental studies using the crustacean stomatogastric nervous system to argue that synaptic and intrinsic currents can vary far more than the output of the circuit in which they are found. These data have significant implications for the mechanisms that maintain stable function over the animal's lifetime, and for the kinds of changes that allow the nervous system to recover function after injury.